[1] Univ Minnesota, Dept Plant & Microbial Biol, 140 Gortner Lab,1479 Gortner Ave, St Paul, MN 55108 USA
[2] Univ Idaho, Dept Biol Sci, 875 Perimeter Dr MS 3051, Moscow, ID 83844 USA
[3] Univ Idaho, Stillinger Herbarium, 875 Perimeter Dr MS 3051, Moscow, ID 83844 USA
[4] Pontificia Univ Catolica Ecuador, Escuela Ciencias Biol, Herbario QCA, Av 12 Octubre 1076 & Roca,Apartado 17-01-2184, Quito, Ecuador