[1] Hokkaido Univ, Grad Sch Med, Dept Dermatol, Sapporo, Hokkaido 0608638, Japan
[2] Univ Dundee, Epithelial Genet Grp, Div Mol Med, Coll Life Sci & Med, Dundee, Scotland
[3] Univ Dundee, Epithelial Genet Grp, Div Mol Med, Coll Dent, Dundee, Scotland
[4] Univ Dundee, Epithelial Genet Grp, Div Mol Med, Coll Nursing, Dundee, Scotland