[1] South African Natl Biodivers Inst, Kirstenbosch Res Ctr, ZA-7735 Claremont, South Africa
[2] Univ Stellenbosch, Dept Bot & Zool, Ctr Invas Biol, ZA-7602 Matieland, South Africa
[3] Univ Adelaide, Sch Biol Sci, North Terrace, Adelaide, SA 5005, Australia
[4] Acad Sci Czech Republic, Dept Invas Ecol, Inst Bot, CZ-25243 Pruhonice, Czech Republic
[5] Charles Univ Prague, Fac Sci, Dept Ecol, CZ-12844 Prague, Czech Republic
[6] UCL, Dept Genet Evolut & Environm, Ctr Biodivers & Environm Res, Gower St, London WC1E 6BT, England
[7] Zool Soc London, Inst Zool, Regents Pk, London NW1 4RY, England