[1] Univ Utrecht, Inst Environm Biol, Lab Palaeobot & Palynol, NL-3584 CD Utrecht, Netherlands
[2] Carl von Ossietzky Univ Oldenburg, Inst Biol Chem & Marine Environm, D-26111 Oldenburg, Germany
[3] Univ Bremen, Ctr Marine Environm Sci, D-28359 Bremen, Germany
[4] Royal Netherlands Inst Sea Res, Dept Marine Biogeochem & Toxicol, NL-1790 AB Den Burg, Netherlands
[5] Univ Utrecht, Dept Earth Sci, NL-3584 CD Utrecht, Netherlands