[1] Univ Texas Austin, Dept Anthropol, Austin, TX 78712 USA
[2] Univ Western Australia, Sch Human Sci, Perth, WA, Australia
[3] Univ Western Australia, Ctr Evolutionary Biol, Sch Biol Sci, Perth, WA, Australia
[4] Kyoto Univ, Primate Res Inst, Inuyama, Aichi, Japan