[1] Shandong Acad Agr Sci, Crop Res Inst, Jinan, Peoples R China
[2] Minist Agr & Rural Affairs, Key Lab Wheat Biol & Genet Improvement North Yell, Jinan, Peoples R China
[3] Natl Engn Lab Wheat & Maize, Jinan, Shandong, Peoples R China
[4] Univ Maryland, Dept Plant Sci & Landscape Architecture, College Pk, MD 20742 USA
[5] Univ Maryland, Inst Biosci & Biotechnol Res, Rockville, MD USA