[1] Cincinnati Childrens Hosp Med Ctr, Visual Syst Grp, Abrahamson Pediat Eye Inst, Div Pediat Ophthalmol, Cincinnati, OH 45229 USA
[2] Kyoto Univ, Grad Sch Pharmaceut Sci, Dept Syst Biol, Sakyo Ku, Kyoto 6068501, Japan
[3] Cincinnati Childrens Hosp Med Ctr, Div Dev Biol, Cincinnati, OH 45229 USA
[4] Univ Cincinnati, Coll Med, Dept Ophthalmol, Cincinnati, OH 45229 USA