[1] Univ Aix Marseille 1, Lab Dynam & Ressources Vegetal EA 2202, UFR DENTES, F-13331 Marseille, France
[2] Univ Aix Marseille 1, SVTE, F-13331 Marseille, France
[3] Ecole Natl Super Chim, F-34296 Montpellier, France
[4] IUT La Rochelle, Dept Appl Biol, Lab SESNAB, F-17026 La Rochelle, France
[5] Univ Ouagadougou, Lab Biol & Physiol Vegetale, Ouagadougou, Burkina Faso