[1] Univ Bath, Milner Ctr Evolut, Dept Biol & Biochem, Bath, England
[2] UCL, Dept Anthropol, London, England
[3] Sun Yat Sen Univ, Sch Ecol, State Key Lab Biocontrol, Shenzhen, Peoples R China
[4] Taif Univ, Coll Sci, Dept Biol, POB 11099, At Taif 21944, Saudi Arabia
[5] Chinese Acad Sci, Northwest Inst Plateau Biol, Xining, Peoples R China
[6] Beijing Normal Univ, Coll Life Sci, Minist Educ, Key Lab Biodivers Sci & Ecol Engn, Beijing, Peoples R China
[7] Univ Debrecen, Dept Evolutionary Zool & Human Biol, HUN REN DE Reprod Strateg Res Grp, Debrecen, Hungary
[8] Univ Debrecen, Debrecen Biodivers Ctr, Debrecen, Hungary