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Free Univ Amsterdam, Fac Med, Dept Anat & Embryol, Inst Neurosci,Grad Sch Neurosci Amsterdam, NL-1081 BT Amsterdam, Netherlands Free Univ Amsterdam, Fac Med, Dept Anat & Embryol, Inst Neurosci,Grad Sch Neurosci Amsterdam, NL-1081 BT Amsterdam, Netherlands