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Sun Yat Sen Univ, Sch Marine Sci, South China Sea Bioresource Exploitat & Utilizat, Guangdong Prov Key Lab Marine Resources & Coastal, Guangzhou 510275, Guangdong, Peoples R China Sun Yat Sen Univ, Sch Marine Sci, South China Sea Bioresource Exploitat & Utilizat, Guangdong Prov Key Lab Marine Resources & Coastal, Guangzhou 510275, Guangdong, Peoples R China