[1] Univ Nacl Jujuy, Fac Ciencias Agr, Ctr Invest & Estudios Diversidad Vegetal CIEDIVE, Alberdi 47, RA-4600 San Salvador De Jujuy, Argentina
[2] UNJu, CONICET, Inst Ecorreg Andinas, Canonigo Gorriti 237, RA-4600 San Salvador De Jujuy, Argentina
[3] Consejo Nacl Invest Cient & Tecn, IBODA, Inst Bot Darwin, Labarden 200,Casilla Correo 22,B1642HYD, Buenos Aires, DF, Argentina
[4] Univ Costa Rica, Escuela Biol, Apdo Postal 11501-2060, San Jose, Costa Rica
[5] Univ Costa Rica, Ctr Invest Biodiversidad & Ecol Trop CIBET, Herbario A Luis Fournier Origgi, Apdo Postal 11501-2060, San Jose, Costa Rica
[6] Leibniz Ctr Agr Landscape Res ZALF, Eberswalder Str 84, D-15374 Muncheberg, Germany
[7] Museo Hist Nat Noel Kempff Mercado, Herbario Oriente USZ, Av Irala 565, Santa Cruz 2489, Bolivia
[8] Univ Bonn, Nees Inst Biodiversitat Pflanzen, Meckenheimer Allee 171, D-53115 Bonn, Germany