[1] Univ Liverpool, Inst Ageing & Chron Dis, Dept Eye & Vis Sci, Liverpool, Merseyside, England
[2] SpheriTech Ltd, Heath Business & Tech Pk, Runcorn, Cheshire, England
[3] Univ Liverpool, Inst Intergrat Biol, Dept Funct & Comparat Genom, Liverpool, Merseyside, England
[4] Royal Liverpool Univ Hosp, St Pauls Eye Unit, Liverpool, Merseyside, England