[1] Univ Sfax, Sfax Fac Sci, Dept Life Sci, Biodivers & Aquat Ecosyst UR 11ES72 Res Unit, Soukra Rd Km 3-5 BP 1171,POB 3000, Sfax, Tunisia
[2] IPEIS, Coastal & Urban Environm Unit, Sfax 3000, Tunisia
[3] Univ Sfax, Fac Sci Sfax, Lab Mat Sci & Environm MESLab, PB 1171, Sfax 3000, Tunisia
[4] Univ Sfax, Fac Sci Sfax, Lab Chem Nat Subst, PB 1171, Sfax 3000, Tunisia
[5] Univ Toulon & Var, Aix Marseille Univ, CNRS INSU, IRD,Mediterranean Inst Oceanog MIO UM 110, F-13288 Marseille, France