[1] Hirosaki Univ, Dept Earth Environm Sci, Hirosaki, Aomori, Japan
[2] Grad Univ Adv Studies, Tachikawa, Tokyo, Japan
[3] Nagasaki Univ, Grad Sch Fisheries Sci & Environm Studies, 1-14 Bunkyo, Nagasaki 8528521, Japan
[4] Japan Agcy Marine Earth Sci & Technol, 2-15 Natsushimacho, Yokosuka, Kanagawa 2370061, Japan
[5] Mie Univ, Climate & Ecosyst Dynam Div, Tsu, Mie 514, Japan
[6] Nagasaki Univ, Fac Fisheries, Nagasaki 852, Japan
[7] Hokkaido Univ, Grad Sch Sci, Sapporo, Hokkaido, Japan