[1] Chinese Acad Sci, Nanjing Inst Geol & Palaeontol, State Key Lab Palaeobiol & Stratig, 39 East Beijing Rd, Nanjing 210008, Jiangsu, Peoples R China
[2] Univ Hong Kong, Dept Earth Sci, Hong Kong, Hong Kong, Peoples R China
[3] Sorbonne Univ, Museum Natl Hist Nat, ISYEB UMR CNRS 7205, Inst Systemat,Evolut,Biodivers,MNHN,UPMC,EPHE,Ent, 57 Rue Cuvier,CP 50, F-75005 Paris, France
[4] Nat Hist Museum, Dept Earth Sci, London SW7 5BD, England
[5] PLA Informat Engn Univ, 62 Kexue Ave, Zhengzhou 450002, Henan, Peoples R China
[6] Chinese Acad Sci, Inst Zool, Key Lab Zool Systemat & Evolut, 1 Beichen West Rd, Beijing 100101, Peoples R China