[1] Rakuno Gakuen Univ, Sch Vet Med, Dept Pharmacol, Hokkaido 0698501, Japan
[2] Gifu Univ, Fac Appl Biol Sci, Pharmacol Lab, Gifu 5011193, Japan
[3] RIKEN, Brain Sci Inst, Mol Neuropathol Grp, Neurogenet Lab, Saitama 3510198, Japan
[4] NIDDK, Bioorgan Chem Lab, Bethesda, MD 20892 USA