[1] Manchester Acad Hlth Sci, NIHR Manchester Musculoskeletal Biomed Res Unit, Manchester, Lancs, England
[2] Univ Manchester, Arthrit Res UK Ctr Genet & Genom, Manchester, Lancs, England
[3] Univ Coll Dublin, UCD Sch Med & Med Sci, Conway Inst, Dublin, Ireland
[4] Univ Leeds, Leeds Teaching Hosp NHS Trust, NIHR Leeds Musculoskeletal Biomed Res Unit, Leeds, W Yorkshire, England
[5] Univ Leeds, Leeds Inst Rheumat & Musculoskeletal Med, Leeds, W Yorkshire, England
[6] Newcastle Univ, NIHR Newcastle Biomed Res Ctr Ageing & Chron Dis, Newcastle Upon Tyne, Tyne & Wear, England
[7] Newcastle Tyne NHS Fdn Trust, Newcastle Upon Tyne, Tyne & Wear, England