[1] Cardiff Univ, Sch Med, Inst Infect & Immun, Cardiff CF14 4XN, S Glam, Wales
[2] Cardiff Univ, Sch Med, Syst Immun Res Inst, Cardiff CF14 4XN, S Glam, Wales
[3] Cardiff Univ, Sch Informat & Comp Sci, Cardiff CF14 4XN, S Glam, Wales
[4] Univ Colorado Denver, Dept Pharmacol, Aurora, CO 80045 USA
[5] Univ Alabama Birmingham, Dept Pathol, Birmingham, AL 35294 USA