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Chinese Ctr Dis Control & Prevent, Natl Inst Parasit Dis, Chinese Ctr Trop Dis Res, WHO Collaborating Ctr Trop Dis,Natl Ctr Int Res Tr, Shanghai 200025, Peoples R China Chinese Ctr Dis Control & Prevent, Natl Inst Parasit Dis, Chinese Ctr Trop Dis Res, WHO Collaborating Ctr Trop Dis,Natl Ctr Int Res Tr, Shanghai 200025, Peoples R China