[1] William Harvey Res Inst, Dept Biochem Pharmacol, London EC1M 6BQ, England
[2] William Harvey Res Inst, Bone & Joint Res Unit, London EC1M 6BQ, England
[3] Natl Inst Adv Ind Sci & Technol, Res Ctr Glycosci, Ibaraki, Japan
[4] Teikyo Univ, Dept Biol Chem, Sagamiko, Kanagawa, Japan