[1] Univ Napoli Parthenope, Ctr Direz, Dipartimento Sci & Tecnol, Isola C4, I-80143 Naples, Italy
[2] Univ Charleston, Grice Marine Lab, 205 Ft Johnson Rd, Charleston, SC 29412 USA
[3] CSIRO Oceans & Atmosphere, Hobart, Tas, Australia
[4] Ctr Southern Hemisphere Oceans Res, Hobart, Tas, Australia
[5] Australian Antarctic Program Partnership, Hobart, Tas, Australia