[1] Univ Michigan, Dept Ecol & Evolutionary Biol, Ann Arbor, MI 48109 USA
[2] Univ Connecticut, Dept Ecol & Evolutionary Biol, Storrs, CT 06269 USA
[3] Washington Univ St Louis, Dept Biol, St Louis, MO 63130 USA
[4] Univ Idaho, Dept Biol Sci, Moscow, ID 83844 USA
[5] Univ Calif Berkeley, Miller Inst Basic Res Sci, Berkeley, CA 94720 USA
[6] Yale Univ, Dept Ecol & Evolutionary Biol, New Haven, CT 06520 USA
[7] Michigan State Univ, Dept Integrat Biol & Ecol, Evolut & Behav Program, Dept Microbiol & Mol Genet, E Lansing, MI 48824 USA
[8] Univ Michigan, Ctr Study Complex Syst, Ann Arbor, MI 48109 USA