[1] Univ Manchester, Arthrit Res UK Ctr Genet & Genom, Ctr Musculoskeletal Res, Manchester M13 9PL, Lancs, England
[2] Cent Manchester Fdn Trust, NIHR Manchester Musculoskeletal Biomed Res Unit, Manchester, Lancs, England
[3] Univ Manchester, Manchester Acad Hlth Sci, Manchester, Lancs, England
[4] Univ Erlangen Nurnberg, Inst Human Genet, Erlangen, Germany
[5] Univ Crete, Dept Internal Med, Lab Mol Med & Human Genet, Iraklion, Greece
[6] Univ Leeds, Leeds Inst Rheumat & Musculoskeletal Med, NIHR Leeds Musculoskeletal Biomed Res Unit, Leeds, W Yorkshire, England
[7] Trinity Coll Dublin, Inst Mol Med, Dept Clin Med, Dublin, Ireland
[8] Adelaide & Meath Hosp, Dept Rheumatol, Dublin, Ireland
[9] Trinity Coll Dublin, Dublin, Ireland
[10] Univ Bath, Royal Natl Hosp Rheumat Dis, Bath BA2 7AY, Avon, England
[11] Univ Bath, Dept Pharm & Pharmacol, Bath BA2 7AY, Avon, England
[12] Univ Roma Tor Vergata, Dept Biomed & Prevent, Rome, Italy
[13] Fdn Santa Lucia IRCCS, Lab Mol Genet UILDM, Rome, Italy
[14] Keele Univ, Inst Sci & Technol Med, Haywood Hosp, Rheumatol Dept,Hlth Serv Res Unit, Stoke On Trent, Staffs, England
[15] Univ Coll Dublin, St Vincents Univ Hosp, UCD Sch Med & Med Sci, Dept Rheumatol, Dublin 2, Ireland
[16] Univ Coll Dublin, Conway Inst Biomol & Biomed Res, Dublin 2, Ireland
[17] Goethe Univ Frankfurt, Div Rheumatol, D-60054 Frankfurt, Germany
[18] Goethe Univ Frankfurt, Fraunhofer IME Project Grp Translat Med & Pharmac, D-60054 Frankfurt, Germany
[19] Cent Manchester Fdn Trust, NIHR Manchester Biomed Res Ctr, Kellgren Ctr Rheumatol, Manchester, Lancs, England
[20] CSIC, Inst Parasitol & Biomed Lopez Neyra, Granada, Spain
[21] Hosp La Princesa, IIS IPrincesa, Dept Rheumatol, Madrid, Spain