[1] Russian Acad Sci, Shirshov Inst Oceanol, 36 Nakhimovskiy Ave, Moscow 117997, Russia
[2] NN Zubov State Oceanog Inst, 6 Kropotkinskiy Lane, Moscow 119034, Russia
[3] Norwegian Inst Water Res, Gaustadalleen 21, N-0345 Oslo, Norway
[4] Russian Acad Sci, AN Severtsov Inst Ecol & Evolut, 33 Leninsky Ave, Moscow 119071, Russia
[5] European Commiss, Joint Res Ctr, Via Enrico Fermi 2749, I-21027 Ispra, VA, Italy
[6] Russian Acad Sci, Far Eastern Branch, VI Ilyichov Pacific Oceanol Inst, 43 Baltic St, Vladivostok 690041, Russia