[1] Chinese Acad Sci, Inst Tibetan Plateau Res, Key Lab Alpine Ecol, Beijing 100101, Peoples R China
[2] Univ Chinese Acad Sci, Beijing 10049, Peoples R China
[3] CAS Ctr Excellence Tibetan Plateau Earth Sci, Beijing 100101, Peoples R China
[4] Bacha Khan Univ Charsadda, Dept Bot, Kp, Pakistan
[5] Friedrich Alexander Univ Erlangen Nuremberg, Inst Geog, Wetterkreuz 15, D-91058 Erlangen, Germany