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Chinese Acad Sci, Ctr Excellence Mol Plant Sci, Inst Plant Physiol & Ecol, Natl Ctr Gene Res,State Key Lab Plant Mol Genet, Shanghai 200233, Peoples R China Chinese Acad Sci, Ctr Excellence Mol Plant Sci, Inst Plant Physiol & Ecol, Natl Ctr Gene Res,State Key Lab Plant Mol Genet, Shanghai 200233, Peoples R China