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Chinese Acad Sci, South China Sea Inst Oceanol, Key Lab Trop Marine Bioresource & Ecol, Guangdong Key Lab Marine Meteria Med, 164 Xingangxi Rd, Guangzhou 510301, Guangdong, Peoples R China Chinese Acad Sci, South China Sea Inst Oceanol, Key Lab Trop Marine Bioresource & Ecol, Guangdong Key Lab Marine Meteria Med, 164 Xingangxi Rd, Guangzhou 510301, Guangdong, Peoples R China