[1] Univ Tolima, Fac Ciencias, Dept Biol, Grp Invest Zool, Calle 42 1B-1, Ibague, Tolima, Colombia
[2] Inst Invest Recursos Biol Alexander von Humboldt, Colecc Biol, Claustro San Agustin, Carrera 8 15-08, Villa De Leyva, Boyaca, Colombia
[3] Univ Estadual Paulista, UNESP, Inst Biociencias, Dept Morfol, Campus Botucatu, BR-18618689 Botucatu, SP, Brazil
[4] Pontificia Univ Javeriana, Unidad Ecol & Sistemat UNESIS, Lab Ictiol, Ciencias Biol, Carrera 7 43-82, Bogota, Colombia
[5] 1822 N Charles St, Belleville, IL 62221 USA