[1] Univ Paris 11, Ecol Systemat & Evolut, UMR CNRS 8079, FR-91405 Orsay, France
[2] Sorbonne Univ, Univ Paris 06, Unite Biol Organismes & Ecosyst Aquat,CNRS,IRD, BOREA,UMR 7208,Museum Natl Hist Nat,Univ Caen Bas, Paris, France
[3] UCL, Dept Genet Evolut & Environm, Ctr Biodivers & Environm Res, London WC1E 6BT, England
[4] Univ Calif Los Angeles, Dept Ecol & Evolutionary Biol, Los Angeles, CA 90095 USA
[5] Univ Calif Los Angeles, Ctr Trop Res, Inst Environm & Sustainabil, Los Angeles, CA 90095 USA
[6] Montpellier SupAgro, INRA, UMR CBGP, IRD, 755 Ave Campus Agropolis,CS 30016, FR-34988 Montferrier Sur Lez, France
[7] Coll William & Mary, Virginia Inst Marine Sci, POB 1346, Gloucester Point, VA 23062 USA