[1] Murdoch Univ, Harry Butler Inst, 90 South St, Murdoch, WA 6150, Australia
[2] Western Australian Museum, Dept Terr Zool, Locked Bag 49, Welshpool Dc, WA 6986, Australia
[3] Ctr Taxon & Morphol, Leibnitz Inst Anal Biodivers Change LIB, Zool Museum Hamburg, Martin Luther King Pl 3, D-20146 Hamburg, Germany
[4] Univ Fed Rio de Janeiro, Univ Brasil, Lab Diversidade Aracnideos, Av Carlos Chagas Filho 373, BR-21941902 Rio De Janeiro, Brazil