[1] Chinese Acad Sci, Inst Zool, Key Lab Zool Systemat & Evolut, 1 Beichen West Rd, Beijing 100101, Peoples R China
[2] Natl Univ Singapore, Dept Biol Sci, 14 Sci Dr 4, Singapore 117543, Singapore
[3] Utah State Univ, Insects Res Unit, USDA ARS Pollinating, Logan, UT 84322 USA
[4] Chinese Acad Sci, Inst Zool, State Key Lab Integrated Pest Management, 1 Beichen West Rd, Beijing 100101, Peoples R China
[5] Univ Chinese Acad Sci, Coll Life Sci, 19 A Yuquan Rd, Beijing 100049, Peoples R China