[1] Queen Mary Univ London, Sch Biol & Chem Sci, London E1 4NS, England
[2] US Geol Survey, Wetland & Aquat Res Ctr, Ft Lauderdale, FL 33314 USA
[3] Panthera, 8 West 40th St,18th Floor, New York, NY 10018 USA
[4] Univ Pretoria, Mammal Res Inst, Dept Zool & Entomol, Lynnwood Rd, ZA-0002 Pretoria, South Africa