[1] Chinese Acad Sci, Key Lab Vertebrate Evolut & Human Origins, Inst Vertebrate Paleontol & Paleoanthropol, Beijing, Peoples R China
[2] China Univ Geosci, Sch Earth Sci & Resources, Beijing, Peoples R China
[3] Chinese Acad Sci, Key Lab Evolutionary Systemat Vertebrates, Beijing, Peoples R China
[4] Univ Colorado, Dinosaur Trackers Res Grp, Denver, CO 80202 USA
[5] Saurierwelt Palaontol Museum, Alte Richt 7, Neumarkt, Germany
[6] Henan Polytech Univ, Collaborat Innovat Ctr Coalbed Methane & Shale Ga, Inst Resources & Environm, Key Lab Biogen Traces & Sedimentary Minerals Hena, Jiaozuo, Henan, Peoples R China
[7] Coll Charleston, Mace Brown Museum Nat Hist, Dept Geol & Environm Geosci, Charleston, SC 29401 USA