[1] Chengdu Ctr China Geol Survey, Chengdu, Sichuan, Peoples R China
[2] Yunnan Univ, Centre Vertebrate Evolutionary Biol, Kunming, Yunnan, Peoples R China
[3] Chinese Acad Sci, Inst Vertebrate Paleontol & Paleoanthropol, Key Lab Vertebrate Evolut & Human Origins, Beijing, Peoples R China
[4] China Univ Geosci Wuhan, Sch Earth Sci, Wuhan, Hubei, Peoples R China
[5] Univ Alberta, Dept Biol Sci, Edmonton, AB, Canada
[6] Philip J Currie Dinosaur Museum, Wembley, AB, Canada