[1] Newcastle Univ, Sch Nat & Environm Sci, Dove Marine Lab, Cullercoats NE30 4PZ, Tyne & Wear, England
[2] Univ Exeter, Coll Life & Environm Sci, Dept Biosci, Hatherly Labs, Exeter EX4 4PS, England
[3] Benthic Solut Ltd, Greengates Way, Hoveton NR12 8ED, Norfolk, England
[4] Newcastle Univ, BioScreening Core Facil, Campus Ageing & Vital, Newcastle Upon Tyne NE4 7RU, England
[5] Newcastle Univ, Sch Nat & Environm Sci, Ridley Bldg, Newcastle Upon Tyne NE1 7RU, England