[1] Wildlife Conservat Soc, Plot 802 Kiwafu Rd,POB 7487, Kampala, Uganda
[2] Univ Queensland, Sch Biol Sci, Brisbane, Qld 4072, Australia
[3] Uganda Wildlife Author, Kampala, Uganda
[4] Wildlife Conservat Soc, Quartier Des Volcans, Goma, DEM REP CONGO
[5] Univ Queensland, Sch Geog Planning & Environm Management, Brisbane, Qld 4072, Australia
[6] Wildlife Conservat Soc, Global Conservat Program, Bronx, NY 10460 USA
[7] Univ London Imperial Coll Sci Technol & Med, Div Ecol & Evolut, Ascot SL5 7PY, Berks, England