[1] Sun Yat Sen Univ, Sch Life Sci, Guangdong Higher Educ Inst, State Key Lab Biocontrol,Key Lab Biodivers Dynam, Guangzhou 510275, Guangdong, Peoples R China
[2] CSIRO Natl Res Collect Australia, Australian Natl Insect Collect, GPO Box 1700, Canberra, ACT 2601, Australia
[3] Chinese Acad Sci, Inst Zool, Key Lab Zool Systemat & Evolut, Beijing 100101, Peoples R China
[4] Chinese Acad Sci, Nanjing Inst Geol & Palaeontol, State Key Lab Palaeobiol & Stratig, 39 East Beijing Rd, Nanjing 210008, Jiangsu, Peoples R China
[5] Chinese Acad Sci, Ctr Excellence Life & Paleoenvironm, 39 East Beijing Rd, Nanjing 210008, Jiangsu, Peoples R China