[1] Chinese Acad Sci, Inst Zool, Key Lab Zool Systemat & Evolut, Beijing 100101, Peoples R China
[2] Chinese Acad Sci, Nanjing Inst Geol & Palaeontol, State Key Lab Palaeobiol & Stratig, 39 East Beijing Rd, Nanjing 210008, Peoples R China
[3] Chinese Acad Sci, Ctr Excellence Life & Paleoenvironm, 39 East Beijing Rd, Nanjing 210008, Peoples R China
[4] Iowa State Univ, Dept Ecol Evolut & Organismal Biol, 2200 Osborn Dr,Bessey Hall, Ames, IA 50011 USA
[5] Univ Gdansk, Dept Invertebrate Zool & Parasitol, Wita Stwosza 59, PL-80308 Gdansk, Poland