[1] Chinese Acad Sci, Northwest Inst Ecoenvironm & Resources, Key Lab Desert & Desertificat, Lanzhou 730000, Peoples R China
[2] South China Normal Univ, Sch Geog, Guangzhou 510631, Peoples R China
[3] Lanzhou Univ, Coll Earth & Environm Sci, Key Lab Western Chinas Environm Syst MOE, Lanzhou 730000, Peoples R China
[4] Univ Helsinki, Environm Change Res Unit, Ecosyst & Environm Res Programme, Fac Biol & Environm Sci, POB 65, Helsinki 00014, Finland
[5] Univ Helsinki, Fac Biol & Environm Sci, Helsinki Inst Sustainabil Sci, POB 65, Helsinki 00014, Finland