[1] Southern Marine Sci & Engn Guangdong Lab Guangzho, Guangzhou 511458, Peoples R China
[2] Chinese Acad Sci, Guangdong Prov Observat & Res Stn Coastal Upwelli, South China Sea Inst Oceanol, State Key Lab Trop Oceanog,Key Lab Sci & Technol, Guangzhou 511458, Peoples R China
[3] Guangdong Ocean Univ, Sch Elect & Informat Engn, Zhanjiang 524025, Peoples R China
[4] State Ocean Adm, South China Sea Environm Monitoring Ctr, Guangzhou 510300, Peoples R China
[5] Chinese Acad Sci, Sanya Inst Oceanol, South China Sea Inst Oceanol, Sanya 572025, Peoples R China