[1] Univ Helsinki, Dept Forest Sci, POB 27, FIN-00014 Helsinki, Finland
[2] Swedish Univ Agr Sci, Dept Forest Ecol & Management, Umea, Sweden
[3] Swedish Univ Agr Sci, Umea Plant Sci Ctr, Dept Forest Genet & Plant Physiol, Umea, Sweden
[4] Norwegian Univ Life Sci NMBU, Fac Environm Sci & Nat Resource Management, As, Norway
[5] Norwegian Inst Bioecon Res NIBIO, Div Forestry & Forest Resources, N-1431 As, Norway
[6] Rockefeller Univ, Program Human Environm, 1230 York Ave, New York, NY 10021 USA