[1] Chinese Acad Sci, Inst Tibetan Plateau Res, Key Lab Alpine Ecol & Biodivers, Beijing 100085, Peoples R China
[2] CAS Ctr Excellence Tibetan Plateau Earth Sci, Beijing 100085, Peoples R China
[3] Cold & Arid Reg Environm & Engn Res Inst, State Key Lab Cryospher Sci, Lanzhou 730000, Peoples R China
[4] Chinese Acad Sci, China Gen Microbiol Culture Collect Ctr, Inst Microbiol, Beijing 100101, Peoples R China