[1] Chinese Acad Sci, South China Bot Garden, Key Lab Vegetat Restorat & Management Degraded Ec, Guangzhou 510650, Guangdong, Peoples R China
[2] Chinese Acad Sci, Inst Subtrop Agr, Key Lab Agroecol Proc Subtrop Reg, Huanjiang Observat & Res Stn Karst Ecosyst, Changsha 410125, Hunan, Peoples R China
[3] Chinese Acad Sci, Inst Geog Sci & Nat Resources Res, Beijing 100101, Peoples R China
[4] Univ Copenhagen, Dept Geosci & Nat Resource Management, DK-1958 Frederiksberg C, Denmark
[5] Chinese Acad Sci, Inst Bot, State Key Lab Vegetat Environm Change, Beijing 100093, Peoples R China
[6] Sinodanish Ctr Educ & Res, DK-8000 Aarhus C, Denmark