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Guizhou Univ, Collaborat Innovat Ctr Mt Ecol & AgroBioengn CICM, Coll Life Sci,Inst Agrobioengn, Minist Educ,Key Lab Plant Resource Conservat & Ge, Guiyang 550025, Guizhou, Peoples R China Guizhou Univ, Collaborat Innovat Ctr Mt Ecol & AgroBioengn CICM, Coll Life Sci,Inst Agrobioengn, Minist Educ,Key Lab Plant Resource Conservat & Ge, Guiyang 550025, Guizhou, Peoples R China