[1] Univ Manchester, Fac Life Sci, Wellcome Trust Ctr Cell Matrix Res, Manchester M13 9PT, Lancs, England
[2] Univ Manchester, Fac Life Sci, Manchester M13 9PT, Lancs, England
[3] Univ Copenhagen, Inst Sports Med, Bispebjerg Hosp, Fac Hlth Sci, Copenhagen, Denmark