[1] Childrens Hosp Philadelphia, Philadelphia, PA 19104 USA
[2] Univ Sydney, Cent Inst Canc Med & Cell Biol, Newtown, Tas, Australia
[3] Univ Penn, Med Ctr, Hemophillia Ctr W Pennsylvania, Pittsburgh, PA USA
[4] Wistar Inst Anat & Biol, Program Immunol, Philadelphia, PA 19104 USA
[5] Howard Hughes Med Inst, Philadelphia, PA USA