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South China Agr Univ, Natl Key Lab Green Pesticide, Guangdong Prov Key Lab Microbial Signals & Dis Con, Integrat Microbiol Res Ctr,Coll Plant Protect, Guangzhou 510642, Peoples R China South China Agr Univ, Natl Key Lab Green Pesticide, Guangdong Prov Key Lab Microbial Signals & Dis Con, Integrat Microbiol Res Ctr,Coll Plant Protect, Guangzhou 510642, Peoples R China