[1] Univ Montpellier 2, CNRS, Inst Sci Evolut ISE M, F-34095 Montpellier 05, France
[2] Univ Hassan II Casablanca, Fac Sci Ain Chock, Lab Ecol Aquat & Environm, Casablanca, Morocco
[3] Tour Valat, Ctr Rech Conservat Zones Humides Mediterraneennes, F-13200 Arles, France
[4] Univ Cadi Ayyad, Fac Polydisciplinaire Safi, Lab Environm Littoral & Marin, Safi, Morocco
[5] Univ Tunis El Manor, Fac Sci Tunis, Dept Biol, Tunis 2092, Tunisia
[6] Univ Manouba, Fac Lettres & Humanites Manouba, UR Biogeog Climatol Appl &Dynam Eros UR99 02 04, Manouba 2010, Tunisia