[1] Natl Ctr Neurol & Psychiat, Natl Inst Neurosci, Dept Anim Models Human Dis, Kodaira, Tokyo 1878502, Japan
[2] Kyoto Univ, Primate Res Inst, Dept Behav & Brain Sci, Inuyama, Aichi 484, Japan
[3] Japan Soc Promot Sci, Tokyo, Japan
[4] Japan Sci & Technol Agcy, Core Res Evolut Sci & Technol, Kawaguchi, Saitama, Japan